you lights are too high to call your setup a low light tank, you should start to dose Excel. yellowing of plant indicate lack of nitrate or Iron, in most cases Nitrate. holes indicate lack of potassium, are you dosing any fertilizer??
there is no real number for how much potassium you should add, plant can handle the high potassium very well. if i have to give you numbers, i would say 20-30ppm total.