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i have collected this plant from many locations in north carolina. in the wild, it grew in various types of substrate, from gravel to mud, and in various types of water, from soft acidic to hard alkaline. i have seen it growing in full sun and in heavy shade.

the one thing that is common in every instance that i have encountered it is that it barely struggles to survive in stagnant water but THRIVES in fast flowing water.

see all the green in the water around me and in my net in this pic? it is all M. umbrosum.
as far as i can tell, as long as you have decent light and ferts, the only thing else that makes a difference is flow.
 
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